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The 1999 book by Bruce Bagemihl, Ph.D. (New York: St. Martin's Press, 1999, Publisher's Weekly "Best Book of 1999"), "Biological Exuberance, Animal Homosexuality and Natural Diversity," states that homosexual and/or transgender activity has been observed in "over 450 species." He states in his book's introduction:
"Any book on homosexuality and transgender in animals is necessarily unfinished, a work in progress. The subject is so vast, the types of behaviors so varied, and the number of species involved so large, as to defy any attempts at comprehensiveness. And the scientific research in this area is only in its infancy: new developments and discoveries are continually being made, and the extent of uncharted and as yet unknowable terrain is so great as to render any attempt at completeness hopelessly premature."
His book states that, as of 1999, homosexual and/or transgender behavior has been observed in the following 478 species:
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